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Cibotium × heleniae

D.D.Palmer (1999)

This name is accepted

Kingdom: Viridiplantae Phylum: Pteridophyta Class/Clade: Polypodiopsida Order: Cyatheales Family: Cibotiaceae Genus: Cibotium

hapu, hāpu‘u

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Description

Key Characters:

Growth Form:

Stems:

Roots:

Leaves:

Sori:

Gametophyte:

Ploidy:

Habitat:

Elevation Range:

Historical Distribution

Images

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Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

O'ahu Endemic
Molokai Endemic
Maui Endemic

Dispersal Agents


Bibliography

Name Published In: Amer. Fern J. 88(4): 150 (1999)

Other References

Palmer 2003:98 (O, parentage: C. chamissoi × C. menziesii); Oppenheimer & Pezzillo 2024:59 (WM); Note: assumed unstabilized hybrid

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Cibotium × heleniae Moanalua Valley, central ridge adjacent to Palmer 831 (C. chamissoi) and Palmer 832 (C. splendens). Preserved_Specimen Palmer, D.D. 830 Oahu BISH 1992-02-13
2 Cibotium × heleniae Between Kalahuapueo and Hanalilolilo Preserved_Specimen Krajina, V.J. 620611029 Molokai BISH 1962-06-11
3 Cibotium × heleniae Waahila Ridge Preserved_Specimen Palmer, D.D. 2255 Oahu BISH 1995-04-23
4 Cibotium × heleniae Aiea Ridge trail Preserved_Specimen Palmer, D.D. 2318 Oahu BISH 1995-08-20
5 Cibotium × heleniae Central ridge of Moanalua Valley Preserved_Specimen Palmer, D.D. 809 Oahu BISH 1992-01-24
6 Cibotium × heleniae Waahila Ridge (St. Louis Heights). On the side of the trail 2/3rds of the way to the summit from the park Preserved_Specimen Palmer, D.D. 845 Oahu BISH 1992-03-01
7 Cibotium × heleniae Poamoho Trail, Koolau Mts Preserved_Specimen Palmer, D.D. 786 Oahu BISH 1992-01-09
8 Cibotium × heleniae Kaala summit bog Preserved_Specimen Palmer, D.D. 738 Oahu BISH 1991-08-22
9 Cibotium × heleniae Aiea, ridge to summit Preserved_Specimen Palmer, D.D. 975 Oahu BISH 1992-08-29
10 Cibotium × heleniae Hawaii Loa ridge Preserved_Specimen Palmer, D.D. 2327 Oahu BISH 1995-09-04
11 Cibotium × heleniae Waahila Ridge (St. Louis Heights). On the side of the trail 2/3rds of the way to the summit from the park Preserved_Specimen Palmer, D.D. 845 Oahu BISH 1992-03-01
12 Cibotium × heleniae Kaala summit bog Preserved_Specimen Palmer, D.D. 738 Oahu BISH 1991-08-22
13 Cibotium × heleniae Hawaii Loa ridge Preserved_Specimen Palmer, D.D. 2328 Oahu BISH 1995-09-04
14 Cibotium × heleniae Waahila Ridge (St. Louis Heights). On the side of the trail 2/3rds of the way to the summit from the park Preserved_Specimen Palmer, D.D. 845 Oahu BISH 1992-03-01
15 Cibotium × heleniae East Maui, near Hana, just W of Wailauki Gulch 26.8N 156.0W Preserved_Specimen Krajina, V.J. 620420000 Maui BISH 1962-04-20
16 Cibotium × heleniae Central ridge of Moanalua Valley Preserved_Specimen Palmer, D.D. 809 Oahu BISH 1992-01-24
17 Cibotium × heleniae Poamoho Trail, Koolau Mts Preserved_Specimen Palmer, D.D. 786 Oahu BISH 1992-01-09
18 Cibotium × heleniae Aiea, ridge to summit Preserved_Specimen Palmer, D.D. 975 Oahu BISH 1992-08-29
19 Cibotium × heleniae Aiea, ridge to summit Preserved_Specimen Palmer, D.D. 975 Oahu BISH 1992-08-29
20 Cibotium × heleniae Hawaii Loa ridge Preserved_Specimen Palmer, D.D. 2327 Oahu BISH 1995-09-04
21 Cibotium × heleniae Poamoho Trail, Koolau Mts Preserved_Specimen Palmer, D.D. 786 Oahu BISH 1992-01-09
22 Cibotium × heleniae Waahila Ridge (St. Louis Heights). On the side of the trail 2/3rds of the way to the summit from the park Preserved_Specimen Palmer, D.D. 845 Oahu BISH 1992-03-01
23 Cibotium × heleniae Aiea, ridge to summit Preserved_Specimen Palmer, D.D. 975 Oahu BISH 1992-08-29
24 Cibotium × heleniae Waahia Ridge. In proximity of both putative parents. Preserved_Specimen Palmer, D.D. 2253 Oahu BISH 1995-04-23
25 Cibotium × heleniae Hawaii Loa ridge Preserved_Specimen Palmer, D.D. 2328 Oahu BISH 1995-09-04
26 Cibotium × heleniae Central ridge of Moanalua Valley Preserved_Specimen Palmer, D.D. 809 Oahu BISH 1992-01-24
27 Cibotium × heleniae Hawaii Loa ridge Preserved_Specimen Palmer, D.D. 2328 Oahu BISH 1995-09-04
28 Cibotium × heleniae Poamoho Trail, Koolau Mts Preserved_Specimen Palmer, D.D. 786 Oahu BISH 1992-01-09
29 Cibotium × heleniae Aiea, ridge to summit Preserved_Specimen Palmer, D.D. 975 Oahu BISH 1992-08-29
30 Cibotium × heleniae Kaala summit bog Preserved_Specimen Palmer, D.D. 738 Oahu BISH 1991-08-22
31 Cibotium × heleniae East Maui, near Hana, just W of Wailauki Gulch 26.8N 156.0W Preserved_Specimen Krajina, V.J. 620420007 Maui BISH 1962-04-20
32 Cibotium × heleniae Kaala summit bog Preserved_Specimen Palmer, D.D. 738 Oahu BISH 1991-08-22
33 Cibotium × heleniae Waialae Nui ridge, 20' down on E side close association with C. chamissoi and C. splendens| Preserved_Specimen Palmer, D.D. 637 Oahu BISH 1991-06-26
34 Cibotium × heleniae Aina Haina Valley, middle ridge, 3/4 way to summit of Koolau Mts In association with C. chamissoi and C. menziesii. Preserved_Specimen Palmer, D.D. 3103 Oahu BISH 1999-03-07
35 Cibotium × heleniae Waahila Ridge, 2/3 way to summit from park side of trail Preserved_Specimen Palmer, D.D. 846 Oahu BISH 1992-03-01
36 Cibotium × heleniae Waiahole Trail base of pali. In close association (10 feet) with both putative ancestors. Preserved_Specimen Palmer, D.D. 806 Oahu BISH 1992-01-20
37 Cibotium × heleniae Halawa Tunnel, side ridge near leeward mouth with mixed population of both putative parents Preserved_Specimen Palmer, D.D. 804 Oahu BISH 1992-01-16
38 Cibotium × heleniae Kaala summit bog Preserved_Specimen Palmer, D.D. 779 Oahu BISH 1991-10-01
39 Cibotium × heleniae Halawa Tunnel, side ridge near leeward mouth with mixed population of both putative parents Preserved_Specimen Palmer, D.D. 804 Oahu BISH 1992-01-16
40 Cibotium × heleniae Waiahole Trail base of pali. In close association (10 feet) with both putative ancestors. Preserved_Specimen Palmer, D.D. 806 Oahu BISH 1992-01-20
41 Cibotium × heleniae Moanalua Valley central ridge in association w/ C. chamissoi & C. splendens. Preserved_Specimen Palmer, D.D. 827 Oahu BISH 1992-02-13
42 Cibotium × heleniae Aina Haina Valley, middle ridge, 3/4 way to summit of Koolau Mts In association with C. chamissoi and C. menziesii. Preserved_Specimen Palmer, D.D. 3103 Oahu BISH 1999-02-22
43 Cibotium × heleniae Aina Haina Valley, middle ridge, 3/4 way to summit of Koolau Mts In association with C. chamissoi and C. menziesii. Preserved_Specimen Palmer, D.D. 3103 Oahu BISH 1999-03-07
44 Cibotium × heleniae Waialae Nui Ridge, 100' down from ridge top Plant in close association with both putative ancestors. Preserved_Specimen Palmer, D.D. 813 Oahu BISH 1992-01-26
45 Cibotium × heleniae Aiea ridge to summit trail side Preserved_Specimen Palmer, D.D. 842 Oahu BISH 1992-02-25
46 Cibotium × heleniae Hawaii Loa ridge near both putative parents Preserved_Specimen Palmer, D.D. 2326 Oahu BISH 1995-09-04
47 Cibotium × heleniae Kaala summit bog Preserved_Specimen Palmer, D.D. 779 Oahu BISH 1991-10-01
48 Cibotium × heleniae Aiea ridge to summit trail side Preserved_Specimen Palmer, D.D. 842 Oahu BISH 1992-02-25
49 Cibotium × heleniae Aiea ridge near both putative parents Preserved_Specimen Palmer, D.D. 2421 Oahu BISH 1996-01-14
50 Cibotium × heleniae Oahu, 1/2 the distance to the summit [presumably Mt. Olympus] side of trail 1/2 way to summit Preserved_Specimen Palmer, D.D. 1154 Oahu BISH 1993-06-20
51 Cibotium × heleniae Manana Ridge trail associated with both putative ancestors (C. menziesii, C. chamissoi) Preserved_Specimen Palmer, D.D. 875 Oahu BISH 1992-06-04
52 Cibotium × heleniae Aiea ridge to summit trail side Preserved_Specimen Palmer, D.D. 842 Oahu BISH 1992-02-25
53 Cibotium × heleniae Aiea ridge trail with both parents Preserved_Specimen Palmer, D.D. 2319 Oahu BISH 1995-11-19
54 Cibotium × heleniae Waiahole Gulch. Along trail between the water tunnels at the base of the pali Collected adjacent to both 601 and 602 (Cibotium chamissoi & C. splendens). Preserved_Specimen Palmer, D.D. 603 Oahu BISH 1991-06-03
55 Cibotium × heleniae Waialae Nui Ridge, 100' down from ridge top Plant in close association with both putative ancestors. Preserved_Specimen Palmer, D.D. 813 Oahu BISH 1992-01-26
56 Cibotium × heleniae Kaala summit bog side of board walk Preserved_Specimen Palmer, D.D. 1163 Oahu BISH 1993-06-21
57 Cibotium × heleniae Waiahole Gulch. Along trail between the water tunnels at the base of the pali Collected adjacent to both 601 and 602 (Cibotium chamissoi & C. splendens). Preserved_Specimen Palmer, D.D. 603 Oahu BISH 1991-06-03
58 Cibotium × heleniae Oahu, 1/2 the distance to the summit [presumably Mt. Olympus] side of trail 1/2 way to summit Preserved_Specimen Palmer, D.D. 1154 Oahu BISH 1993-06-20
59 Cibotium × heleniae Halawa Tunnel, side ridge near leeward mouth with mixed population of both putative parents Preserved_Specimen Palmer, D.D. 804 Oahu BISH 1992-01-16
60 Cibotium × heleniae Waiahole Trail base of pali. In close association (10 feet) with both putative ancestors. Preserved_Specimen Palmer, D.D. 806 Oahu BISH 1992-01-20
61 Cibotium × heleniae Kaala summit bog side of board walk Preserved_Specimen Palmer, D.D. 1163 Oahu BISH 1993-06-21
62 Cibotium × heleniae Kaala summit bog side of board walk Preserved_Specimen Palmer, D.D. 1163 Oahu BISH 1993-06-21
63 Cibotium × heleniae Poamoho Trail in association with both ancestors Preserved_Specimen Palmer, D.D. 1186 Oahu BISH 1993-08-08
64 Cibotium × heleniae Waiahole Gulch. Along trail between the water tunnels at the base of the pali Collected adjacent to both 601 and 602 (Cibotium chamissoi & C. splendens). Preserved_Specimen Palmer, D.D. 603 Oahu BISH 1991-06-03
65 Cibotium × heleniae Waialae Nui ridge, 20' down on E side close association with C. chamissoi and C. splendens| Preserved_Specimen Palmer, D.D. 637 Oahu BISH 1991-06-26
66 Cibotium × heleniae Oahu, 1/2 the distance to the summit [presumably Mt. Olympus] side of trail 1/2 way to summit Preserved_Specimen Palmer, D.D. 1154 Oahu BISH 1993-06-20
67 Cibotium × heleniae Manana Ridge trail associated with both putative ancestors (C. menziesii, C. chamissoi) Preserved_Specimen Palmer, D.D. 875 Oahu BISH 1992-06-04
68 Cibotium × heleniae Mt. Kaala summit bog Preserved_Specimen Palmer, D.D. 732 Oahu BISH 1991-08-22
69 Cibotium × heleniae W Maui, Wailuku District, Waihee Valley, back of valley on plateau Northing: 2313353; Easting: 753356 Rare. Growing in shady forest with C. menziesii and C. glaucum. In Metrosideros Wet Forest. Preserved_Specimen Oppenheimer, H.L. 61608 Maui BISH 2016-06-20
70 Cibotium × heleniae Hawaii Loa ridge near both putative parents Preserved_Specimen Palmer, D.D. 2326 Oahu BISH 1995-09-04
71 Cibotium × heleniae Kipapa Trail, 3/4+ mi from refuge boundary, S side of trail UTM 610757.083E, 2374326.05N| Metrosideros, Acacia, Dicranopteris; wet lowland forest. Uncommon to moderately common on and off trail; parental types common. Preserved_Specimen Reynolds, L.S. 140 Oahu BISH 2006-06-13
72 Cibotium × heleniae Mt. Kaala summit bog Preserved_Specimen Palmer, D.D. 732 Oahu BISH 1991-08-22
73 Cibotium × heleniae Kaala summit bog Preserved_Specimen Palmer, D.D. 779 Oahu BISH 1991-10-01
74 Cibotium × heleniae Waahila Ridge, 2/3 way to summit from park side of trail Preserved_Specimen Palmer, D.D. 846 Oahu BISH 1992-03-01
75 Cibotium × heleniae Waialae Nui ridge, 20' down on E side close association with C. chamissoi and C. splendens| Preserved_Specimen Palmer, D.D. 637 Oahu BISH 1991-06-26
76 Cibotium × heleniae Moanalua Valley, central ridge adjacent to Palmer 831 (C. chamissoi) and Palmer 832 (C. splendens). Preserved_Specimen Palmer, D.D. 830 Oahu BISH 1992-02-13
77 Cibotium × heleniae Waialae Nui Ridge, 100' down from ridge top Plant in close association with both putative ancestors. Preserved_Specimen Palmer, D.D. 813 Oahu BISH 1992-01-26
78 Cibotium × heleniae Mt. Kaala summit bog Preserved_Specimen Palmer, D.D. 732 Oahu BISH 1991-08-22
79 Cibotium × heleniae Mt. Kaala; road up north slope; Waianae Mts Metrosideros forest near summit. Preserved_Specimen Little Jr., E.L. 31301 Oahu BISH 1978-10-18
80 Cibotium × heleniae Halawa Tunnel, side ridge near leeward mouth with mixed population of both putative parents Preserved_Specimen Palmer, D.D. 804 Oahu BISH 1992-01-16
81 Cibotium × heleniae Manana Ridge trail associated with both putative ancestors (C. menziesii, C. chamissoi) Preserved_Specimen Palmer, D.D. 875 Oahu BISH 1992-06-04
82 Cibotium × heleniae Aiea ridge trail with both parents Preserved_Specimen Palmer, D.D. 2319 Oahu BISH 1995-11-19
83 Cibotium × heleniae Aina Haina Valley, middle ridge, 3/4 way to summit of Koolau Mts In association with C. chamissoi and C. menziesii. Preserved_Specimen Palmer, D.D. 3103 Oahu BISH 1999-03-07
84 Cibotium × heleniae Kaala, summit associated w/ C. chamissoi, C. splendens, and C. glaucum. Preserved_Specimen Palmer, D.D. 866 Oahu BISH 1991-04-08
85 Cibotium × heleniae Mt. Kaala; road up north slope; Waianae Mts Metrosideros forest near summit. Preserved_Specimen Little Jr., E.L. 31301 Oahu BISH 1978-10-18
86 Cibotium × heleniae Waialae Nui ridge, 20' down on E side close association with C. chamissoi and C. splendens| Preserved_Specimen Palmer, D.D. 637 Oahu BISH 1991-06-26
87 Cibotium × heleniae Moanalua Valley central ridge in association w/ C. chamissoi & C. splendens. Preserved_Specimen Palmer, D.D. 827 Oahu BISH 1992-02-13
88 Cibotium × heleniae Aiea ridge near both putative parents Preserved_Specimen Palmer, D.D. 2421 Oahu BISH 1996-01-14
89 Cibotium × heleniae Kaala summit bog Preserved_Specimen Palmer, D.D. 779 Oahu BISH 1991-10-01
90 Cibotium × heleniae W Maui, Wailuku District, Waihee Valley, back of valley on plateau Northing: 2313353; Easting: 753356 Rare. Growing in shady forest with C. menziesii and C. glaucum. In Metrosideros Wet Forest. Preserved_Specimen Oppenheimer, H.L. 61608 Maui BISH 2016-06-20
91 Cibotium × heleniae Kipapa Trail, 3/4+ mi from refuge boundary, S side of trail UTM 610757.083E, 2374326.05N| Metrosideros, Acacia, Dicranopteris; wet lowland forest. Uncommon to moderately common on and off trail; parental types common. Preserved_Specimen Reynolds, L.S. 140 Oahu BISH 2006-06-13
92 Cibotium × heleniae Moanalua Valley, central ridge adjacent to Palmer 831 (C. chamissoi) and Palmer 832 (C. splendens). Preserved_Specimen Palmer, D.D. 830 Oahu BISH 1992-02-13
93 Cibotium × heleniae Waiahole Gulch, about 800 ft along trail between water tunnels at base of pali Collected adjacent to both 601 and 602 (Cibotium chamissoi & C. splendens). Preserved_Specimen Palmer, D.D. 603 Oahu BISH 1991-06-03
94 Cibotium × heleniae Waahila Ridge, 2/3 way to summit from park side of trail Preserved_Specimen Palmer, D.D. 846 Oahu BISH 1992-03-01
95 Cibotium × heleniae Kaala, summit associated w/ C. chamissoi, C. splendens, and C. glaucum. Preserved_Specimen Palmer, D.D. 866 Oahu BISH 1991-04-08
96 Cibotium × heleniae Halawa Tunnel, side ridge near leeward mouth with mixed population of both putative parents Preserved_Specimen Palmer, D.D. 804 Oahu BISH 1992-01-16
97 Cibotium × heleniae Waiahole Gulch, about 800 ft along trail between water tunnels at base of pali Collected adjacent to both 601 and 602 (Cibotium chamissoi & C. splendens). Preserved_Specimen Palmer, D.D. 603 Oahu BISH 1991-06-03
98 Cibotium × heleniae Waialae Nui Ridge, 100' down from ridge top Plant in close association with both putative ancestors. Preserved_Specimen Palmer, D.D. 813 Oahu BISH 1992-01-26
99 Cibotium × heleniae Mt. Kaala summit bog Preserved_Specimen Palmer, D.D. 732 Oahu BISH 1991-08-22
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